Publications

If not already available Open Access, PDFs of publications are available freely by request. Simply send a message via the Contact form and include the paper(s) you would like.

70. Heads, S.W. 2024. A second chrysopine lacewing from Mexican amber (Insecta: Neuroptera: Chrysopidae). Kentiana 6: 1–6. https://doi.org/10.21900/j.kentiana.n6.1807

69. Skejo, J., Kasalo, N., Thomas, M.J. & Heads, S.W. 2024. A new long-winged pygmy grasshopper in Eocene Baltic amber raises questions about the evolution of reduced tegmenula in Tetrigidae (Orthoptera). Journal of Orthoptera Research 33: 21–26. https://doi.org/10.3897/jor.33.105144

68. Jiménez-Mejías, P., Manzano, S., Gowda, V., Frank Thorsten-Krell, et al., incl. Heads, S.W. among 1,559 additional authors. 2024. Protecting stable biological nomenclatural systems enables universal communication: a collective international appeal BioScience 74(7): 467–472. https://doi.org/10.1093/biosci/biae043

67. Hedlund, T., Thomas, M.J. & Heads, S.W. 2023. The first record of Phaulotettix ablusus from the United States (Orthoptera: Acrididae: Melanoplinae). Entomological News 130(5): 463–465. https://doi.org/10.3157/021.130.0508

66. Dietrich, C.H., Williams, C.J. & Heads, S.W. 2023. A new species and new combination in the planthopper genus Cheiloceps Uhler (Hemiptera: Fulgoroidea: Issidae) from Miocene Dominican amber. Kentiana 4: 1–8. https://doi.org/10.21900/j.kentiana.n4.1401

65. Heads, S.W., Thomas, M.J., Hedlund, T.J. & Wang, Y. 2023. A new fossil katydid of the genus Arethaea Stål (Orthoptera: Tettigoniidae) with exceptionally preserved internal organs from the Eocene Green River Formation of Colorado. Palaeoentomology 6(3): 268–277. https://doi.org/10.11646/palaeoentomology.6.3.10

64. Heads, S.W., Kolata, D.R. & Guensburg, T.E. 2023. On the type material of the Upper Ordovician mitrate “carpoid” Diamphidiocystis drepanon Kolata & Guensburg (Echinodermata: Stylophora). Kentiana 3: 1–2. https://doi.org/10.21900/j.kentiana.n3.1179

63. Heads, S.W. & Taylor, S.J. 2022. First record of a fossil coral (Anthozoa) from Pleistocene deposits at Terra Ceia, North Carolina. Kentiana 2: 1–4. https://doi.org/10.21900/j.kentiana.n2.1160

62. Heads, S.W., Thuy, B. & Tamarri, J. 2022. Two new species of Archelcana Sharov (Orthoptera: Elcanidae) from the Lower Jurassic of Luxembourg. Kentiana 1: 1–13. https://doi.org/10.21900/j.kentiana.n1.1121

61. Wang, B., Shi, G., Xu, C., Spicer, R.A., Perrichot, V., Schmidt, A.R., Feldberg, K., Heinrichs, J., Chény, C., Pang, H., Liu, X., Gao, T., Wang, Z., Ślipiński, A., Solórzano-Kraemer, M., Heads, S.W., Thomas, M.J., Sadowski, E.M., Szwedo, J., Azar, D., Nel, A., Liu, C., Chen, J., Zhang, Q., Luo, C., Yu, T., Zheng, D., Zhang, H. & Engel, M.S. 2021. The mid-Miocene Zhangpu biota reveals an outstandingly rich rainforest biome in East Asia. Science Advances 7: 1–7, eabg0625. https://doi.org/10.1126/sciadv.abg0625

60. Barling, N.T., Heads, S.W. & Martill, D.M. 2021. Addendum: Behavioural impacts on the taphonomy of dragonflies (Odonata) from the Lower Cretaceous Crato Formation, Brazil. Palaeoentomology 4: 203. https://doi.org/10.11646/palaeoentomology.4.3.3

59. Barling, N.T., Heads, S.W. & Martill, D.M. 2021. Behavioural impacts on the taphonomy of dragonflies and damselflies (Odonata) from the Lower Cretaceous Crato Formation, Brazil. Palaeoentomology 4: 141–155. https://doi.org/10.11646/palaeoentomology.4.2.3

58. Swanson, D.R., Heads, S.W., Taylor, S.J. & Wang, Y. 2021. A new remarkably preserved fossil assassin bug (Insecta: Heteroptera: Reduviidae) from the Eocene Green River Formation of Colorado. Papers in Palaeontology 7(3): 1459–1478. https://doi.org/10.1002/spp2.1349

57. Swanson, D.R. & Heads, S.W. 2020. Rescuing a lost damsel: a revised identity for the extinct true bug Miocoris fagi (Heteroptera). Palaeoentomology 3: 597–606. https://doi.org/10.11646/palaeoentomology.3.6.11

56. Barling, N.T., Martill, D.M., & Heads, S.W. 2020. A geochemical model for the preservation of insects in the Crato Formation (Lower Cretaceous) of Brazil. Cretaceous Research 116: 104608. https://doi.org/10.1016/j.cretres.2020.104608

55. Xu, C., Fang, Y., Heads, S.W., Zhang, Q., Jarzembowski, E.A. & Wang, B. 2020. The first Orthoptera (Insecta) from the Triassic of China. Alcheringa 44: 93–98. https://doi.org/10.1080/03115518.2019.1662485

54. Thomas, M.J., Skejo, J. & Heads, S.W. 2019. The last batrachideine of Europe: A new genus and species of pygmy grasshopper (Orthoptera: Tetrigidae) from Eocene Baltic amber. Zootaxa 4686: 435–445. https://doi.org/10.11646/zootaxa.4686.3.9

53. Heads, S.W., Thomas, M.J. & Wang, Y. 2018. A new genus and species of Elcanidae (Insecta: Orthoptera) from Cretaceous Burmese amber. Zootaxa 4527: 575–580. https://doi.org/10.11646/zootaxa.4527.4.8

52. Baena-Bejarano, N., Heads, S.W. & Taylor, S.J. 2018. Comments on the neglected nymphs of mud crickets in the genus Mirhipipteryx (Caelifera: Tridactyloidea: Ripipterygidae). Zootaxa 4486: 180–188. https://doi.org/10.11646/zootaxa.4486.2.7

51. Heads, S.W. 2018. On the status of Araripegryllus romualdoi (Insecta: Orthoptera: Grylloidea) from the Cretaceous Santana Formation of Brazil. Zootaxa 4486: 83–84. https://doi.org/10.11646/zootaxa.4486.1.6

50. Fang, Y., Muscente, A.D., Heads, S.W., Wang, B. & Xiao, S. 2018. The earliest Elcanidae (Insecta, Orthoptera) from the Upper Triassic of North America. Journal of Paleontology 92: 1028–1034. https://doi.org/10.1017/jpa.2018.20

49. Robinson, J.L., Thomas, M.J. & Heads, S.W. 2018. A fossil caddisfly (Insecta, Trichoptera) from the Eocene of Colorado. Zootaxa 4403: 389–394. https://doi.org/10.11646/zootaxa.4403.2.10

48. Baena-Bejarano, N. & Heads, S.W. 2018. First records of Dentridactylinae (Orthoptera: Tridactylidae) from Colombia, with description of two new species of Dentridactylus Günther. Zootaxa 4402: 575–584. https://doi.org/10.11646/zootaxa.4402.3.10

47. Fang, Y., Heads, S.W., Wang, H., Zhang, H. & Wang, B. 2018. The first Archelcaninae (Orthoptera: Elcanidae) from the Cretaceous Jehol Biota of Liaoning, China. Cretaceous Research 86: 129–134. https://doi.org/10.1016/j.cretres.2018.02.008

46. Makarkin, V.N., Wedmann, S. & Heads, S.W. 2018. A systematic reappraisal of Araripeneuridae (Neuroptera: Myrmeleontoidea), with description of new species from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 84: 600–621. https://doi.org/10.1016/j.cretres.2017.12.018

45. Heads, S.W., Miller, A.N. & Crane, J.L. 2017. On the name of the oldest fossil mushroom. Mycological Progress, 16: 11–12. https://doi.org/10.1007/s11557-017-1355-4

44. Heads, S.W., Miller, A.N., Crane, J.L., Thomas, M.J., Ruffatto, D.M., Methven, A.S., Raudabaugh, D.B. & Wang, Y. 2017. The oldest fossil mushroom. PLoS ONE 12(6): e0178327, 1–6. https://doi.org/10.1371/journal.pone.0178327

43. Makarkin, V.N., Heads, S.W. & Wedmann, S. 2017. Taxonomic study of the Cretaceous lacewing family Babinskaiidae (Neuroptera: Myrmeleontoidea: Nymphidoidae), with description of new taxa. Cretaceous Research 78: 149–160. https://doi.org/10.1016/j.cretres.2017.06.007

42. Swanson, D.R., Taylor, S.J. & Heads, S.W. 2017. Nine true bugs (Heteroptera) newly-discovered in Illinois. Zootaxa 4269: 571–585. https://doi.org/10.11646/zootaxa.4269.4.10

41. Heads, S.W. & Hollier, J. 2016. The type specimen and generic placement of Tridactylus galla Saussure, 1895 (Orthoptera: Caelifera: Tridactylidae). Zootaxa 4193: 195–196. https://doi.org/10.11646/zootaxa.4193.1.12

40. Jepson, J.E. & Heads, S.W. 2016. New Megaloptera (Insecta: Neuropterida) from the Lower Cretaceous Crato Formation of Brazil. Zootaxa 4098: 134–144. https://doi.org/10.11646/zootaxa.4098.1.5

39. Song, H., Amédégnato, C., Cigliano, M.M., Desutter-Grandcolas, L., Heads, S.W., Huang, Y., Otte, D. & Whiting, M.F. 2015. 300 million years of diversification: elucidating the patterns of orthopteran evolution based on comprehensive taxon and gene sampling. Cladistics 31: 621–651. https://doi.org/10.1111/cla.12116

38. Hollier, J. & Heads, S.W. 2015. An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 6: The Raphidophoroidea, Stenopelmatoidea and Tettigonioidea. Revue Suisse de Zoology 112: 307–323. https://doi.org/10.5281/zenodo.30002

37. Baena-Bejarano, N.J. & Heads, S.W. 2015. Three new species of the genus Ripipteryx from Colombia (Orthoptera, Ripipterygidae). ZooKeys 502: 129–143. https://doi.org/10.3897/zookeys.502.8871

36. Barling, N., Martill, D.M., Heads, S.W. & Gallien, F. 2015. High fidelity preservation of fossil insects from the Crato Formation (Lower Cretaceous) of Brazil. Cretaceous Research 52: 605–622. https://doi.org/10.1016/j.cretres.2014.05.007

35. Heads, S.W., Thomas, M.J. & Wang, Y. 2014. A remarkable new pygmy grasshopper (Orthoptera, Tetrigidae) in Miocene amber from the Dominican Republic. ZooKeys 429: 87–100. https://doi.org/10.3897/zookeys.429.8020

34. Hollier, J. & Heads, S.W. 2014. The type specimens of Orthoptera described by American entomologists in the Muséum d’histoire naturelle de Genève. Revue Suisse de Zoology 121: 63–76.

33. Heads, S.W. & Thomas, M.J. 2014. Rediscovery of the Milton Sanderson Collection at the Illinois Natural History Survey, University of Illinois at Urbana-Champaign. Society for the Preservation of Natural History Collections (SPNHC) Connection 28: 32–33.

32. Barling, N., Heads, S.W. & Martill, D.M. 2013. A new parasitoid wasp (Hymenoptera: Chalcidoidea) from the Lower Cretaceous Crato Formation of Brazil: the first Mesozoic Pteromalidae. Cretaceous Research 45: 258–264. https://doi.org/10.1016/j.cretres.2013.05.001

31. Hollier, J., Bruckner, H. & Heads, S.W. 2013. An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 5: Grylloidea. Revue Suisse de Zoology 120: 445–535.

30. Heads, S.W. & Wang, Y. 2013. First fossil record of Melanoplus differentialis (Orthoptera: Acrididae: Melanoplinae). Entomological News 123: 33–37. https://doi.org/10.3157/021.123.0108

29. Jepson, J.E., Heads, S.W., Makarkin, V.N. & Ren, D. 2013. New fossil mantidflies (Insecta, Neuroptera, Mantispidae) from the Mesozoic of northeastern China. Palaeontology 56: 603–613. https://doi.org/10.1111/pala.12005

28. Heads, S.W., Penney, D. & Green, D.I. 2012. A new species of the cricket genus Proanaxipha in Miocene amber from the Dominican Republic (Orthoptera, Gryllidae, Pentacentrinae). ZooKeys 229: 111–118. https://doi.org/10.3897/zookeys.229.3678

27. Pérez-de la Fuente, R., Heads, S.W., Hinojosa-Díaz, I.A. & Engel, M.S. 2012. The first record of Protogryllinae from the Jurassic of India (Orthoptera: Grylloidea). Journal of the Kansas Entomological Society 85: 53–58. https://doi.org/10.2317/JKES111103.1

26. Hollier, J. & Heads, S.W. 2012. An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 1: Tridactyloidea. Revue Suisse de Zoology 119: 149–160.

25. Rodríguez, F. & Heads, S.W. 2012. Erratum: Fernando Rodríguez & Sam W. Heads (2012) New mole crickets of the genus Scapteriscus Scudder (Orthoptera: Gryllotalpidae: Scapteriscinae) from Colombia. Zootaxa 3337: 68. https://doi.org/10.11646/zootaxa.3337.1.3

24. Rodríguez, F. & Heads, S.W. 2012. New mole crickets of the genus Scapteriscus Scudder (Orthoptera: Gryllotalpidae: Scapteriscinae) from Colombia. Zootaxa 3282: 61–68. https://doi.org/10.11646/zootaxa.3282.1.4

23. Heads, S.W. & Taylor, S.J. 2012. A new species of Ripipteryx from Belize with a key to the species of the Scrofulosa Group (Orthoptera, Ripipterygidae). ZooKeys 169: 1–8. https://doi.org/10.3897/zookeys.169.2531

22. Heads, S.W. & Leuzinger, L. 2011. On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera: Ensifera). ZooKeys 77: 17–30. https://doi.org/10.3897/zookeys.77.769

21. Heads, S.W. 2010. New Tridactyloidea in Miocene amber from the Dominican Republic (Orthoptera: Caelifera). Annales de la Société Entomologique de France (N.S.) 46: 204–210. https://doi.org/10.1080/00379271.2010.10697659

20. Heads, S.W. 2010. A new species of Ripipteryx from the Ecuadorian Andes (Orthoptera: Tridactyloidea: Ripipterygidae). Zootaxa 2476: 23–29. https://doi.org/10.11646/zootaxa.2476.1.3

19. Heads, S.W. 2010. The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae) 20 million years of troglobiomorphosis or exaptation in the dark? Zoological Journal of the Linnean Society 158: 56–65. https://doi.org/10.1111/j.1096-3642.2009.00587.x

18. Heads, S.W. & Maehr, M.D. 2009. Proposed conservation of the name Gastrimargus Saussure, 1884 (Orthoptera: Acrididae: Oedipodinae) threatened by an unused senior homonym. Zootaxa 2268: 65–68. https://doi.org/10.11646/zootaxa.2268.1.6

17. Heads, S.W. 2009. A new pygmy mole cricket in Cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia 26: 75–82.

16. Heads, S.W. 2009. New pygmy grasshoppers in Miocene amber from the Dominican Republic (Orthoptera: Tetrigidae). Denisia 26: 69–74.

15. Delclòs, X., Nel, A., Azar, D., Bechly, G., Dunlop, J.A., Engel, M.S. & Heads, S.W. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 247: 353–381. https://doi.org/10.1127/0077-7749/2008/0247-0353

14. Heads, S.W. & Chesmore, D. 2008. New records of the slender ground-hopper Tetrix subulata (Orthoptera: Tetrigidae) from Yorkshire. British Journal of Entomology & Natural History 21: 243–246.

13. Heads, S.W. 2008. A new species of Yuripopovia (Coleorrhyncha: Progonocimicidae) from the Early Cretaceous of the Isle of Wight. British Journal of Entomology & Natural History 21: 247–254.

12. Heads, S.W. & Lakin, C. 2008. A new katydid of the genus Phoebolampta from St. Maarten (Orthoptera: Tettigoniidae: Phaneropterinae). Polski Pismo Entomologiczne 77: 256–266.

11. Heads, S.W. 2008. The first fossil Proscopiidae (Insecta, Orthoptera, Eumastacoidea) with comments on the historical biogeography and evolution of the family. Palaeontology 51: 499–507. https://doi.org/10.1111/j.1475-4983.2008.00756.x

10. Heads, S.W., Martill, D.M. & Loveridge, R.F. 2008. Palaeoentomological paradise: the Cretaceous Crato Formation of Brazil. Antenna 32: 91–98.

9. Martill, D.M., Bechly, G. & Heads, S.W. 2007. Appendix: species list for the Crato Formation, pp. 582–607 in Martill, D.M., Bechly, G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp. [book chapter]

8. Martins-Neto, R.G., Heads, S.W. & Bechly, G. 2007. Neuropterida: snakeflies, dobsonflies and lacewings, pp. 328–340 in Martill, D.M., Bechly, G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp. [book chapter]

7. Menon, F., Heads, S.W. & Szwedo, J. 2007. Cicadomorpha: cicadas and relatives, pp. 283–297 in Martill, D.M., Bechly, G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp. [book chapter]

6. Heads, S.W. & Martins-Neto, R.G. 2007. Orthopterida: grasshoppers, crickets, locusts and stick insects, pp. 265–283 in Martill, D.M., Bechly, G. & Loveridge, R.F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press, xvi + 625 pp. [book chapter]

5. Martill, D.M. & Heads, S.W. 2007. Araripe GeoPark is a palaeontological paradise of international significance, but is the scientific community being excluded? Geoscientist 17: 20–24.

4. Heads, S.W. 2006. A new caddisfly larval case (Insecta, Trichoptera) from the Lower Cretaceous Vectis Formation (Wealden Group) of the Isle of Wight, southern England. Proceedings of the Geologists’ Association 117: 307–310. https://doi.org/10.1016/S0016-7878(06)80037-4

3. Menon, F. & Heads, S.W. 2005. New species of Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) 357: 1–11.

2. Menon, F., Heads, S.W. & Martill, D.M. 2005. New Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 26: 837–844. https://doi.org/10.1016/j.cretres.2005.05.005

1. Heads, S.W., Martill, D.M. & Loveridge, R.F. 2005. An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology 48: 1409–1417. http://doi.org/10.1111/j.1475-4983.2005.00514.x